Original paper

A new lycopsid from the Lower Carboniferous of Nanjing, South China

Wang, Yi; Xu, Hong-He; Jiang, Qing


Biformistrobus wui gen. et sp. nov. is described from the uppermost part of the Wutung Formation (Tournaisian, Early Carboniferous) of Kongshan, Nanjing, South China, as a lycopsid bearing a stem up to 17 mm wide, spirally arranged, narrowly obovate leaf bases, and linear to lanceolate leaves. Stems are multi-dichotomously branched, bearing two different strobili on distal tips, but the anatomical features and in situ spores are unknown. In light of the single-veined microphylls, relatively thin axes, adaxially attached sporangium on the sporophylls, and dimorphic strobili, Biformistrobus probably represents either a part of a monoecious, herbaceous lycopsid or a portion of distal branching systems from a heterosporous, arborescent lycopsid. Biformistrobus is probably endemic in the Early Carboniferous flora of South China.


lycopsidbiformistrobuscarboniferousevolutionwutung formationkongshansouth china