Original paper

Early Miocene ants (Hymenoptera, Formicidae) from Radoboj, Croatia

Dlussky, Gennady M.; Putyatina, Tatyana S.


The ant assemblage of Radoboj (Croatia) described by Heer (1849, 1867) is considered the richest known Miocene assemblage of Europe. However, Heer's data can no longer be used for analysis of the palaeontological history of ants, because they are strongly outdated and require a revision. Such a revision was the purpose of our study. We found in collections of three museums of Austria (Universalmuseum Joanneum in Graz, Geologische Bundesanstalt, and Naturhistoricshes Museum in Wien) a total of 537 compression fossils of ants from Radoboj, 459 of which were identified earlier by Heer. We designated the holotypes, lectotypes and neotypes for 54 of the 62 species described by Heer, and subsequently compared the other specimens to these types. As a result, we have identified 350 specimens to subfamily and 309 specimens to species. We re-described 23 species originally described by Heer (1849, 1867) and two species described by Mayr (1867). One genus and eight species are described as new; 27 species and varieties described by Heer are synonymized. The taxonomic placement of eight species described by Heer remains unclear. As a result of our revision, the known assemblage of Radoboj includes 33 species of 15 genera and five subfamilies. The assemblage of Radoboj is especially similar at the subfamily level to the assemblage of Bembridge, UK (Late Eocene) and Stavropol, RF (Middle Miocene).


croatiaearly mioceneformicidaeradobojrevision and new taxa