Original paper

New and poorly known Aptian Acrioceratidae (Acrioceratidae, Ammonoidea) from Cassis - Roquefort-la-Bédoule (Bouches-du-Rhône, France)

Frau, Camille; Bulot, Luc G.; Delanoy, Gérard


Brainaella marcoulinense gen. nov. et sp. nov. is described on the basis of new material from the Upper Aptian of the reference succession of Cassis - Roquefort-la-Bédoule (Bouches-du-Rhône, France). The morphological features of this new taxon suggest affinities with the late Early Aptian Toxoceratoides rochi Casey, a poorly understood species revised in the present contribution. The study of these two taxa sheds new light on the taxonomic status, content and evolution of the long-debated families Helicancylidae Hyatt and Acrioceratidae Vermeulen.