Original paper

Phytogeographic and climatic implications of Permian woods discovered in Oman (Arabian Peninsula)

Berthelin, Martine;Vozenin-Serra, Colette; Broutin, Jean


Many permineralized woods are found in the Permian Gharif Formation (Sultanate of Oman), associated with palynomorphs, impressions-compressions and isolated fertile structures. The here studied silicified woods are assigned to four distinct gymnospermous taxa:

Baieroxylon implexum (Zimmermann) Greguss 1961, from the Permian of Western Europe, related to Ginkgophyta.
Trigonomyelon omanense n. sp., belonging to a genus previously known from the Permian of India and Brazil, probably assignable to Coniferophyta.
Dadoxylon (Eristophyton) callixyloides n. sp., showing affinities with Dadoxylon (Eristophyton) ibericum from the Permian of Rio Viar (Southwestern of Spain, Vozenin-Serra et al. 1991), Dadoxylon (cf. Eristophyton) from the Tazekka Formation (Morocco, Chalot-Prat & Galtier 1989), and with Cathaysioxylon khmerinum (Vozenin-Serra 1978) from the Kazanian of Cambodia reminiscent of Callixylon. This taxon can probably be assigned to Pteridospermophyta.
Prototaxoxylon gharifense n. sp., which is very similar to the secondary wood of Prototaxopitys americana (Milanez & Dolianiti) Krausel & Dolianiti 1958, from the Permian of Brazil, a putative Ginkgophyta.

All these woods display inconspicuous irregularly distributed growth rings, excluding a marked seasonality, and suggesting a warm and humid climate with slightly dryer periods. The mixed composition of the xyloflora and the macro- and microflora (a mixture of Gondwanan, Euramerican and Cathaysian elements) demonstrate that Oman was located at a tropical to subequatorial paleolatitude during the middle Permian (Wordian).


Le gisement du Permien moyen de la Formation de Gharif (Sultanat d’Oman) a fourni divers bois silicifies trouvés en association avec des palynomorphes et des empreintes (feuilles et organes fertiles isoles). Ces structures ont éte attribuées a quatres genres distincts de Gymnospermes:

Baieroxylon implexum (Zimmermann) Greguss 1961, déja décrit dans le Permien de l’Europe de l’Ouest, rapporte à une Ginkgophyte.
Trigonomyelon omanense n. sp., genre bien connu dans le Permien gondwanien (Inde et Brésil), structure attribuable en toute probabilité à une Conifàrophyte.
Dadoxylon (Eristophyton) callixyloides n. sp. proche de Dadoxylon (Eristophyton) ibericum provenant du Permien du Rio Viar (Sud-Ouest de l’Espagne, Vozenin et al. 1991), de la Formation Tazekka (Maroc, Chalot-Prat & Galtier 1989) et du Cathaysioxylon khmerinum (Vozenin-Serra 1978) du Kazanien cambodgien qui presente aussi une filiation évidente avec le genre Callixylon. Ce taxon pourrait probablement appartenir à une Pteridospermaphyte.
Prototaxoxylon gharifense n.sp. bois secondaire du Protaxopitys americana (Milanez & Dolianiti ) Krausel & Dolianiti 1958, du Permien du Brésil, probable Ginkgophyte.


Fossil woodsPermianArabian Peninsulapaleoclimatologypaleogeography