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Frank Licher:

Revision der Gattung Typosyllis Langerhans, 1879 (Polychaeta: Syllidae)

Morphologie, Taxonomie und Phylogenie

[A revision of Typosyllis Langerhans, 1879 (Polychaeta: Syllidae)]

1999. 336 Seiten, 121 Abbildungen, 24 Tabellen, 21x30cm, 1490 g
Language: Deutsch

(Abhandlungen der Senckenbergischen Naturforschenden Gesellschaft, Band 551)

ISBN 978-3-510-61006-8, brosch., price: 50.00 €

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Contents

Inhaltsverzeichnis top ↑

Einleitung 2
Material und Methode 3
Material 3
Methoden 4
Lichtmikroskopie 4
Rasterelektronenmikroskopie 5
Literaturangaben 5
Cladistische Analyse 6
Abkürzungen 6
Biologie und Verbreitung 7
Lebensweise und Habitat 7
Morphologie 7
Reproduktion 10

Epigamie 11
Schizogamie 11
Viviparie 12
Hermaphroditismus 12
Externe Gestation 12
Laterale Knospung 13
Architomie 13
Steuerung des Fortpflanzungsprozesses 13
Verbreitung der Syllidae 15
Taxonomischer Teil 17
Zur Systematik der Syllidae 17
Die traditionelle Systematik der Syllidae 17
Phylogenie der Syllidae 17
Zur Systematik der Gattung Typosyllis 20
Der "Syllis-Komplex" 20
Phylogenie innerhalb des "Syllis-Komplexes" 21
Grundmuster von Typosyllis 25
Revision der Gattung Typosyllis LANGERHANS, 1879 25
Diagnose 25
Morphologische Merkmale und ihre phylogenetische Bedeutung 27
Bestimmungsschlüssel 31
Beschreibung der distinkten Arten 36
Typosyllis LANGERHANS, 1879 36
nigricirris-Komplex 36
Typosyllis aciculigrossa (SAN MARTIN, 1990) comb. nov. 36
Typosyllis nigricirris (GRUBE, 1863) 37
Typosyllis parateinopterona (HARTMAN & FAUCHALD, 1971) nom. nov. 39
Typosyllis tegulum HARTMAN & FAUCHALD, 1971 41
rosea-Komplex 43
Typosyllis maryae (SAN MARTIN, 1992) comb. nov. 43
Typosyllis rosea (LANGERHANS, 1879) comb. nov. 44
mexicana-Komplex 45
Typosyllis beneliahuae (CAMPOY & ALQUEZAR, 1982) comb. nov., emend. 47
Typosyllis magna (WESTHEIDE, 1974) comb. nov., stat. nov. 49
Typosyllis mexicana (RIOJA, 1960) comb. nov. 51
cornuta-Komplex 53
Typosyllis alosae (SAN MARTIN, 1992) 53
Typosyllis anops (EHLERS, 1897) comb. nov. 54
Typosyllis cornuta (RATHKE, 1843) 57
caeca- Komplex 64
Typosyllis caeca (KATZMANN, 1973) comb. nov. 64
Typosyllis hyperioni (DORSEY & PHILLIPS, 1987) comb. nov. 66
cerina-Komplex 68
Typosyllis botosaneanui HARTMANN-SCHRÖDER, 1973 68
Typosyllis broomensis HARTMANN-SCHRÖDER, 1979 70
Typosyllis cerina (GRUBE, 1878) comb. nov. 72
Typosyllis garciai (CAMPOY, 1982) comb. nov. 74
Typosyllis hyllebergi sp. nov. 76
Typosyllis mouretanica sp. nov. 78
Typosyllis silkeae sp. nov. 81
Typosyllis yallingupensis HARTMANN-SCHRÖDER, 1982 84
parturiens-Komplex 86
Typosyllis golfonovensis HARTMANN-SCHRÖDER, 1962 86
Typosyllis heronislandensis HARTMANN-SCHRÖDER,1991 88
Typosyllis parturiens (HASWELL,1920) 89
Typosyllis patriciae HARTMANN-SCHRÖDER, 1981 90
variegata-Komplex 92
Typosyllis aciculata TREADWELL, 1919 92
Typosyllis bella CHAMBERLIN, 1919 94
Typosyllis macrodentata HARTMANN-SCHRÖDER, 1982 96
Typosyllis remanei HARTMANN-SCHRÖDER, 1960 98
Typosyllis taiwanensis sp. nov. 100
Typosyllis variegata (GRUBE, 1860) 101
Typosyllis warrnamboolensis HARTMANN-SCHRÖDER, 1987 108
Typosyllis westheidei (SAN MARTIN, 1984) comb. nov. 111
corallicola-Komplex 112
Typosyllis benguellana (DAY, 1963) 114
Typosyllis bifurcata HARTMANN-SCHRÖDER, 1980 116
Typosyllis corallicola (VERRILL, 1900) 116
Typosyllis monilata IMAJMA, 1966 119
Typosyllis nipponica IMAJMA, 1966 121
Typosllis setoensis IMAJMA, 1966 124
prolifera-Komplex 125
Typosllis busseltonensis HARTMANN-SCHRÖDER, 1982 125
Typosyllis gerlachi HARTMANN-SCHRÖDER, 1960 127
Typosyllis hawaiiensis HARTMANN-SCHRÖDER, 1965 emend. 129
Typosyllis kerguelensis AVERINCEV, 1972 131
Typosyllis microoculata HARTMANN-SCHRÖDER, 1965 133
Typosyllis prolifera (KROHN, 1852) 135
Typosyllis schulzi HARTMANN-SCHRÖDER, 1960 140
Typosyllis tyrrhena LICHER & KUPER, 1998 140
Typosyllis vivipara (KROHN, 1869) 142
pulvinata-Komplex 145
Typosyllis albanyensis HARTMANN-SCHRÖDER, 1984 145
Typosyllis edensis HARTMANN-SCHRÖDER, 1989 147
Typosyllis erikae HARTMANN-SCHRÖDER, 1981 149
Typosyllis horrocksensis HARTMANN-SCHRÖDER, 1981 151
Typosyllis kabilica (BEN-ELIAHU, 1977) comb. nov., stat. nov. 152
Typosyllis nuchalis HARTMANN-SCHRÖDER, 1960 154
Typosyllis pharynxcircumfusata HARTMANN-SCHRÖDER, 1979 156
Typosyllis pulvinata LANGERHANS, 1881 158
Typosyllis truncata (HASWELL, 1920) 160
luteoides-Komplex 162
Typosyllis bifida HARTMANN-SCHRÖDER, 1986 162
Typosyllis luteoides HARTMANN-SCHRÖDER, 1962 163
Typosyllis raygeorgei HARTMANN-SCHRÖDER, 1979 165
torquata-Komplex 169
Typosyllis cruzi (NUNEZ & SAN MARTIN, 1991) comb. nov. 169
Typosyllis curticirris HARTMANN-SCHRÖDER, 1981 171
Typosyllis gerundensis ALOS & CAMPOY, 1981 171
Typosyllis glarearia WESTHEIDE, 1974 175
Typosyllis lutea HARTMANN-SCHRÖDER, 1960 177
Typosyllis riojai (SAN MARTIN, 1990) comb. nov. 178
Typosyllis torquata (MARION & BOBRETZKY, 1875) 180
Typosyllis typica (MOORE, 1909) comb. nov. 183
magnipectinis-Komplex 185
Typosyllis magnipectinis (STORCH, 1967) 185
Typosyllis ortizi (SAN MARTIN, 1992) 186
armillaris-Komplex 189
Typosyllis armillaris (O.F. MÜLLER, 1776) 189
Typosyllis crassicirrata TREADWELL, 1925 emend. 196
Typosyllis ehlersioides MARENZELLER, 1890 199
Typosyllis hyalina (GRUBE, 1863) 199
Typosyllis krohni (EHLERS, 1864) emend. 205
Typosyllis neglecta (GRUBE, 1870) comb. nov. 207
Typosyllis stellaepolaris HARTMANN-SCHRÖDER, 1993 209
Typosyllis valida (GRUBE, 1857) comb. nov. 211
Typosyllis violacea (GRUBE, 1870) comb. nov. 215
amica-Komplex 217
Typosyllis amica (QUATREFAGES, 1866) 217
Typosyllis elongata (JOHNSON, 1901) 219
Typosyllis ferrani (AL6S & SAN MARTIN, 1987) comb. nov. 221
prolixa-Komplex 223
Typosyllis adamantens (TREADWELL, 1914) 223
Typosyllis dayi HARTMANN-SCHRÖDER, 1974 225
Typosyllis magdalena (WESENBERG-LUND, 1962) 227
Typosyllis pectinans (HASWELL, 1920) 229
Typosyllis pigmentata (CHAMBERLIN, 1919) 233
Typosyllis prolixa (EHLERS, 1901) 235
vittata-Komplex 237
Typosyllis dentata HARTMANN-SCHRÖDER, 1960 237
Typosyllis farallonensis BLAKE & WALTON, 1977 239
Typosyllis fasciata (MALMGREN, 1867) 241
Typosyllis flaccida (GRUBE, 1878) comb. nov. 246
Typosyllis lunaris IMAITMA, 1966 246
Typosyllis okadai (FAUVEL, 1934) 248
Typosyllis vittata (GRUBE, 1840) 250
Typosyllis incertae sedis 253
Typosyllis alternata (MOORE, 1908) 253
Typosyllis anoculata HARTMANN-SCHRÖDER, 1962 255
Typosyllis antarctica AVERINCEV, 1972 257
Typosyllis barbata (SAN MARTIN, 1992) 257
Typosyllis danieli (SAN MARTIN, 1992) 258
Typosyllis filiclentata HARTMANN-SCHRÖDER, 1962 261
Typosyllis heterochaeta (MOORE, 1909) comb. nov. 262
Typosyllis heterocirrata HARTMANN-SCHRÖDER, 1960 emend. 264
Typosyllis japonica (IMAJMA, 1966) comb. nov. 265
Typosyllis nigrescens (GRUBE, 1878) comb. nov. 267
Typosyllis pharobroomensis HARTMANN-SCHRÖDER, 1979 269
Distinkte Arten anderer Taxa der Syllidae 270
Amblyosyllis GRUBE, 1857 270
Amblyosyllis formosa (CLAPAREDE, 1863) 270
Autolytus GRUBE, 1851 272
Autolytus prolifer O.F. MÜLLER, 1788 272
Branchiosyllis EHLERS, 1887 272
Branchiosyllis bathyalis (KIRKEGAARD, 1995) comb. nov. 272
Branchiosyllis exilis (GRAVIER, 1900) 272
Branchiosyllis maculata (IMAJIMA, 1966) comb. nov. 274
Branchiosyllis salina (HARTMANN-SCHRÖDER, 1959) comb. nov. 274
Branchiosyllis verrucalosa (AUGENER, 1913) comb. nov. 274
Brania QUATREFAGES, 1866 275
Brania pusilla (DUJARDIN, 1851) 275
Dentatisyllis PERKINS, 1981 275
Dentatisyllis carolinne (DAY, 1973) 275
Dentatisyllis hongkongensis DING, LICHER & WESTHEIDE, 1998 275
Dentatisyllis inflata (MARENZELLER, 1879) comb. nov. 275
Dentatisyllis junoyi LOPEZ GARCIA & SAN MARTIN, 1992 277
Dentatisyllis kiaorensis (HARTMANN-SCHRÖDER, 1992) comb. nov. 277
Dentatisyllis mangalis RUSSEIL, 1995 277
Dentatisyllis mortoni DING, LICHER & WESTHEIDE, 1998 277
Dentatisyllis uebelackerae DING, LICHER & WESTHEIDE, 1998 278
Ehlersia QUATREFAGES, 1865 278
Ehlersia ferrugina LANGERHANS, 1881 278
Eusyllis MALMGREN, 1867 279
Eusyllis blomstrandi MALMGREN, 1867 279
Eusyllis fragilis (WEBSTER, 1879) 279
Exogone ORSTED, 1845 279
Exogone naidina ORSTED, 1845 279
Haplosyllides AUGENER, 1924 279
Haplosyllides floridana AUGENER, 1924 279
Haplosyllis LANGERHANS, 1879 280
Haplosyllis agelas UEBELACKER, 1982 280
Haplosyllis anthagorgicola UTINOMI, 1956 280
Haplosyllis bisetosa HARTMANN-SCHRÖDER, 1960 280
Haplosyllis cephalata VERRILL, 1900 280
Haplosyllis chamaeleon LAUBIER, 1960 280
Haplosyllis cratericola (BUZHINSKAJA, 1980) comb. nov. 280
Haplosyllis depressa AUGENER, 1913 280
Haplosyllis dollfusi (FAUVEL, 1933) 280
Haplosyllis spongicola (GRUBE, 1855) 281
Haplosyllis spongiphila (VERRTLL, 1885) comb. nov. 283
Haplosyllis trifalcata DAY, 1960 284
Haplosyllis xeniaecola HARTMANN-SCHRÖDER, 1993 284
Odontosyllis CLAPAREDE, 1863 284
Odontosyllis (?) corruscans (HASWELL, 1886) comb. nov.,
Species inquirenda 284
Odontosyllis cucullata (M'INTOSH, 1908) 284
Odontosyllis fulgurans (AUDOUIN & MILNE EDWARDS, 1833) 284
Odontosyllis gibba CLAPAREDE, 1863 285
Odontosyllis polycera (SCHMARDA, 1861) 285
Parasphaerosyllis MONRO, 1937 285
Parasphaerosyllis uschakovi (CHLEBOVITSCH, 1959) 285
Pionosyllis MALMGREN, 1867 285
Pionosyllis compacta MALMGREN, 1867 285
Pionosyllis divaricata (KEFERSTEIN, 1862) 285
Pionosyllis longisetosa (HARTMANN-SCHRÖDER, 1990) comb. nov. 286
Pionosyllis pulligera (KROHN, 1852) 286
Pseudobrania SAN MARTIN, 1984 286
Pseudobrania clavata (CLAPAREDE, 1863) 286
Pseudobrania limbata (CLAPAREDE, 1868) 287
Pseudosyllis GRUBE, 1863 287
Pseudosyllis brevipennis GRUBE, 1863 287
Syllides ORSTED, 1845 288
Syllides longocirrata (ORSTED, 1845) 288
Syllis SAVIGNY, 1812 288
Syllis brasiliensis M'INTOSH, 1885 288
Syllis brevicirrata M'INTOSH,1908 288
Syllis brevicirris HANSEN, 1882 288
Syllis cirrita LEE & RHO, 1994 288
Syllis curticirris MONRO, 1937 289
Syllis elongata DAY, 1949 289
Syllis gracilis GRUBE, 1840 289
Syllis longosegmentata GRUBE, 1857 291
Syllis magellanica AUGENER, 1918 292
Syllis monilaris SAVIGNY, 1812 292
Syllis nidrosiensis BIDENKAP, 1907 292
Syllis onkylochaeta HARTMANN-SCHRÖDER, 199l 292
Syllis ramosa M'INTOSH, 1879 292
Syllis robertianae M'INTOSH,1885 293
Syllis sclerolaema EHLERS, 19O1 293

Syllis umbricolor GRUBE, 1878 293
Trypanosyllis CLAPAREDE, 1864 293
Trypanosyllis complanata (TREADWELL, 19O1) comb. nov. 293
Trypanosyllis gigantea (M'INTOSH, 1885) 294
Trypanosyllis latifrons (GRUBE, 1857) 294
Trypanosyllis taeninaeformis (HASWELL, 1886) 295
Trypanosyllis zebra (GRUBE, 1860) 295
Xenosyllis MARION & BOBRETZKY, 1875 296
Xenosyllis scabra (EHLERS, 1864) 296
Syllinae incertae sedis 297
"Typosyllis" attenuata (KNOX, 1960) species inquirenda 297
"Odontosyllis glandulosa AUGENER, 1913 297
"Typosyllis heterosetosa HARTMANN-SCHRÖDER, 199l 298
Invalide Bezeichnungen 300
Nomina dubia 300
Chaetosyllis oerstedi MALMGREN, 1867 300
Ehlersia nepiotoca CAULLERY & MESNIL, 1916 300
Gnathosyllis diplodonta SCHMARDA, 1861 300
Haplosyllis oligochaeta pontica CZERNIAVSKY, 1881 300
Haplosyllis palpata VERRILL, 1900 300
Ioida pontica CZERNIAVSKY, 1881 302
Langerhansia (Syllis) biocula CZERNIAVSKY, 1881 302
Langerhansia (Syllis) valida CZERNIAVSKY, 1881 302
Langerhansia (Syllis) volida paucisetosa CZERNIAVSKY, 1881 302
Nereis blainvillei DELLE CHIAJE, 1828 302
Nereis cirrhigera VIVIANI, 1805 302
Nereis incisa FABRICIUS, 1780 302
Nereis monoceros DALYELL, 1853 303
Nereis mucronata VIVIANI, 1805 303
Nereis noctiluca LINNAEUS, 1761 303
Nereis rudolphii DELLE CHIAJE, 1841 303
Nereis tethycola DELLE CHIAJE, 1825 303
Nereis tiedemanni DELLE CHIAJE, 1828 303
Nereisyllis ornata BLAINVILLLE, 1828 303
Perikoea disingii KINBERG, 1866 303
Pionosyllis lucida CHAMBERLIN, 1919 304
Syllis abyssicola EHLERS, 1874 304
Syllis bacilligera CLAPAREDE, 1868 304
Syllis breviarticulata GRUBE, 1857 304
Syllis brevicollis EHLERS, 1874 304
Syllis brevis SCHMARDA, 1861 304

Syllis caeca MONRO, 1933 304
Syllis californica KINBERG, 1866 305
Syllis ciliata MECZNIKOW, 1865 305
Syllis erythropis GRUBE, 1878 305
Syllis eximia MALM, 1874 305
Syllis fissipara KROHN, 1852 305
Syllis fusicornis SCHMARDA, 1861 305
SNllis jugalaris VERRILL, 1900 305
Syllis Lycochaetus GRUBE, 1878 306
Syllis macroceras GRUBE, 1857 306
Syllis macroura SCHMARDA, 1861 306
Syllis macalosa MILNE EDWARDS, 1845 306
Syllis moniliformis SAVIGNY, 1822 306
Syllis mytilorum STUDER, 1889 306
Syllis normannica CLAPAREDE, 1863 306
Syllis notocera EHLERS, 1905 306
Syllis obscura GRUBE, 1857 306
Syllis pallida VERRILL, 1875 307
Syllis punctulata HASWELL, 1920 307
Syllis quadrifasciata FISCHLI, 1900 307
Syllis remex CHAMBERLIN, 1919 307
Syllis sexoculata EHLERS, 1864 307

Syllis singalisetis GRUBE, 1878 307
Syllis solida GRUBE, 1878 308
Syllis stenura BLANCHARD, 1849 308
Syllis zonata GRUBE, 1857 308
Syllis (Ehlersia) exigua VERRILL, 1900 308
Syllis (Typosyllis) annularis VERRILL, 1900 308

Syllis (Typosllis) diplomorpha VERRILL, 1900 308

Syllis (Typosyllis) filiformis RULLIER, 1972 308
Typosyllis taprobanensis WILLEY, 1905 309

Nomina nuda 309
Dentatisyllis brevicirra 309
Haplosyllis maderensis 309
Haplosyllis oligochaeta massiliensis 309
Syllis brevicirra 309
Syllis coenobita 309
Syllis densa 309
Syllis elegans 309
Syllis macrocephala 310
Syllis nigra 310
Syllis striata 310
Syllis vancouveri 310
Andere nicht valide Nomina 310
Syllis atlantica 310
Syllis atlantica lineata 310
Verfügbares Typenmaterial 310
Cladistische Analyse 319
Programm 319

Artenkomplexe in Typosyllis 319
Für die cladistische Analyse ausgewählte Merkmale 321
Wahl der Außengruppe 321
Matrix und Cladogrammberechnung 323
Diskussion 324
Danksagung 328
Literatur 329