Original paper

Revision of Burmochloritis Godwin-Austen, 1920 in Southeast Asia (Gastropoda: Stylommatophora: Camaenidae)

Páll-Gergely, Barna; Gojšina, Vukašin; Neubert, Eike

Abstract

This paper revises the Southeast Asian Camaenidae that are characterized by a relatively large (>20 mm) shell with multiple reddish spiral bands on the body whorl and a finely pitted protoconch. The anatomy of 2 species is known, Burmochloritis kengtungensis Godwin-Austen, 1920 and Helix massiei L. Morlet, 1891, the latter from this study. Since both are similar to each other and differ from all other camaenids, we include both in the genus Burmochloritis Godwin-Austen, 1920. All additional species with such traits are known from empty shells only. They are provisionally also placed in Burmochloritis, although further information will be necessary to confirm their taxonomic position. Five new species are described from the collection of the Muséum national d’Histoire naturelle (Paris, France): Burmochloritis laotica Páll-Gergely & Neubert n. sp., B. muratovi Páll-Gergely n. sp., B. nordsiecki Páll-Gergely & Neubert n. sp., B. paya Páll-Gergely n. sp., and B. payella Páll-Gergely n. sp. Helix (Chloritis) marimberti Bavay & Dautzenberg, 1900 is a junior synonym of Burmochloritis luctativa (Mabille, 1889). Helix (Chloritis) marimberti var. carinata Bavay & Dautzenberg, 1909, which has been treated as a synonym of Helix marimberti, is here elevated to species level. Since it is a junior homonym of Helix carinata Link, 1807, the replacement name Burmochloritis laocaiensis Páll-Gergely, nom. nov., is proposed. With the current revision, Burmochloritis comprises 15 species and ranges through eastern Myanmar, northern Laos, and northern Vietnam, and there is a single species known from southern Laos. Chloritis anserina and C. theobaldi, which were formally included in Burmochloritis, are moved to the genus Bouchetcamaena Thach, 2018 because they lack multiple reddish bands.

Keywords

Laos • Vietnam • Myanmar • anatomy • shell • taxonomy • new species.