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An unusual tetrapod assemblage from the latest Triassic of the South Eifel region (Rhineland-Palatinate, Germany)

Numberger-Thuy, Lea D.; Delsate, Dominique; Hanker, Pia; Kronfeld, Andrea; Brockmann, Marc A.; Thuy, Ben

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Neues Jahrbuch für Geologie und Paläontologie - Abhandlungen (2026)

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publication en ligne: Mar 30, 2026
manuscrit accepté: Feb 25, 2026
manuscrit reçu: Jan 5, 2026

DOI: 10.1127/njgpa/1307

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Abstract

Tetrapod vertebrates underwent a major turnover by the end of the Triassic, yet their fossil record from strata close to the Triassic-Jurassic boundary is sparse. Here, we describe new tetrapod fossils from the uppermost Triassic Exter Formation of the recently discovered Irrel section in the south Eifel area, Rhineland-Palatinate, Germany. The material consists of isolated teeth and bones from a richly fossiliferous, conglomeratic bonebed, palynologically dated to the upper middle Rhaetian, and deposited in an intertidal channel. The assemblage is remarkably diverse and includes the assumed thalattosaur Pachystropheus, kuehneosaurid archosauromorphs, theropod dinosaurs, pterosaurs, and late-surviving phytosaurs, plagiosaurs and aetosaurs, the latter representing the youngest known to date. The material furthermore includes osteoderms and a fragmentary tooth of an unknown archosaur, possibly a small crocodylomorph, and a fragmentary tooth of an unknown lepidosauromorph or archosauriform with deep longitudinal grooves assumedly for venom delivery. The assemblage is among the most diverse from the latest Triassic and adds to the sparse Rhaetian fossil record of terrestrial tetrapods in general, and more specifically theropod dinosaurs and pterosaurs.

Mots-clefs

Exter Formation • Irrel • aetosaur • theropod • Rhaetian